Professor Emerita Mary-Lou Pardue, an influential faculty member in the MIT Department of Biology, died on June 1. She was 90.Early in her career, Pardue developed a technique called in situ hybridization with her PhD advisor, Joseph Gall, which allows researchers to localize genes on chromosomes. This led to many discoveries, including critical advancements in developmental biology, our understanding of embryonic development, and the structure of chromosomes. She also studied the remarkably complex way organisms respond to stress, such as heat shock, and discovered how telomeres, the ends of chromosomes, in…